Publications

Ferreira HRS, Champagnon J, Alves JA & Lok T (in press) Relationship between wintering site and survival in a migratory waterbird using different migration routes. Oecologia. 

Lok T, van der Geest M, Bom RA, de Goeij P, Piersma T & Bouten W (2023) Prey ingestion rates revealed by back-mounted accelerometers in Eurasian spoonbills. Animal Biotelemetry 11: 5. Open Access.

Reneerkens J, Versluijs TSL, Piersma T, Alves JA, Boorman M, Corse C, Gilg O, Hallgrimsson GT, Lang J, Loos B, Ntiamoa-Baidu Y, Nuoh AA, Potts PM, ten Horn J & Lok T (2020). Low fitness at low latitudes: wintering in the tropics increases migratory delays and mortality rates in an arctic-breeding shorebird. Journal of Animal Ecology 89: 691-703. PDF

Chan Y-C, Tibbitts TL, Lok T, Hassell C, Peng H-B, Ma Z, Zhang Z & Piersma T. (2019) Filling knowledge gaps in a threatened shorebird flyway through satellite tracking.  Journal of Applied Ecology 56: 2305-2315. PDF

Lok T, Hassell CJ, Piersma T, Pradel R & Gimenez O (2019). Accounting for heterogeneity when estimating stopover duration, timing and population size of red knots in Bohai Bay. Ecology and Evolution 9: 6176-6188. Open Access

van der Geest M, van der Lely JAC, van Gils JA, Piersma T & Lok T (2019). Density-dependent growth of bivalves dominating the intertidal zone of Banc d'Arguin, Mauritania: importance of feeding mode, habitat and season. Marine Ecology Progress Series 610: 51-63. PDF

Jouta J, de Goeij P, Lok T, Velilla E, Camphuysen CJ, Leopold M, van der Veer HW, Olff H, Overdijk O & Piersma T (2018). Unexpected diet preferences of Eurasian Spoonbills in the Dutch Wadden Sea: Spoonbills mainly feed on small fish instead of shrimp. Journal of Ornithology 159: 839-849. PDF

Oudman T, de Goeij P, Piersma T & Lok T (2017). Colony-breeding Eurasian Spoonbills in The Netherlands: local limits to population growth with expansion into new areas. Ardea 105: 113-124. PDF

Lok T, Veldhoen L, Overdijk O, Tinbergen JM & Piersma T (2017). An age-dependent fitness cost of migration? Old trans-Saharan migrating spoonbills breed later than those staying in Europe, and late breeders have lower recruitment. Journal of Animal Ecology 86: 998-1009. PDF

van den Hout P, Piersma T, ten Horn J, Spaans B & Lok T (2017). Individual shifts toward safety explain age-related foraging distribution in a gregarious shorebird. Behavioral Ecology 28: 419-428. PDF

Van Gils JA, Lisovski S, Lok T, Meissner W, Ozarowska A, de Fouw J, Rakhimberdiev E, Soloviev MY, Piersma T & Klaassen M (2016) Body shrinkage due to Arctic warming reduces red knot fitness in tropical wintering range. Science 352: 819-821. PDF
 
Conklin JR, Lok T, Melville DS, Riegen AC, Schuckard R, Piersma T & Battley PF (2016) Declining adult survival of New Zealand Bar-tailed Godwits during 2005–2012 despite apparent population stability. Emu 116: 147-157. PDF
 
Piersma T, Lok T, Chen Y, Hassell CJ, Yang H-Y, Boyle A, Slaymaker M, Chan Y-C, Melville DS, Zhang Z-W & Ma Z (2016) Simultaneous declines in summer survival of three shorebird species signals a flyway at risk. Journal of Applied Ecology 53: 479-490. PDF
 
Lok T, Overdijk O & Piersma T (2015) The cost of migration: spoonbills suffer higher mortality during trans-Saharan spring migrations only. Biology Letters 11: 20140944. PDF
 
Lok T, Overdijk O & Piersma T (2014) Interpreting variation in growth of Eurasian Spoonbill chicks: disentangling the effects of age, sex and environment. Ardea 102: 181-194. PDF
 
El-Hacen EM, Piersma T, Jouta J, Overdijk O & Lok T (2014) Seasonal variation in the diet of Spoonbill chicks in the Wadden Sea: a stable isotopes approach. Journal of Ornithology 155: 611-619. PDF
 
Lok T, Overdijk O, Tinbergen JM & Piersma T (2013) Seasonal variation in density dependence in age-specific survival of a long-distance migrant. Ecology 94: 2358-2369. PDF
 
El-Hacen EM, Overdijk O, Lok T, Olff H & Piersma T (2013) Home range, habitat selection, and foraging rhythm in Mauritanian spoonbills (Platalea leucorodia balsaci): a satellite tracking study. Waterbirds 36: 277-286. PDF
 
van Gils JA, van der Geest M, Leyrer J, Oudman T, Lok T, Onrust J, de Fouw J, van der Heide T, van den Hout PJ, Spaans B, Dekinga A, Brugge M & Piersma T (2013) Toxin constraint explains diet choice, survival and population dynamics in a molluscivore shorebird. Proceedings of the Royal Society B-Biological Sciences 280: 20130861. PDF
 
Leyrer J, Lok T, Brugge M, Spaans B, Sandercock BK & Piersma T (2013) Mortality within the annual cycle: seasonal survival patterns in Afro-Siberian red knots. Journal of Ornithology 154: 933-943. PDF
 
Lok T, Overdijk O & Piersma T (2013) Migration tendency delays distributional response to differential survival prospects along a flyway. American Naturalist 181: 520-531. PDF
 
Piersma T, Van der Velde M, El-Hacen HM, Lok T, Overdijk O (2012) Molecular verification of the subspecies status of the Mauritanian spoonbill Platalea leucorodia balsaci. Ardea 100: 131-136. PDF
 
Leyrer J, Lok T, Brugge M, Dekinga A, Spaans B, Van Gils JA, Sandercock BK & Piersma T (2012) Small-scale demographic structure suggests preemptive behavior in a flocking shorebird. Behavioral Ecology 23: 1226-1233. PDF
 
Lok T, Overdijk O, Tinbergen JM & Piersma T (2011) The paradox of spoonbill migration: most birds travel to where survival rates are lowest. Animal Behaviour 82: 837-844. PDF
 
van de Pol M, Ens BJ, Heg D, Brouwer L, Krol J, Maier M, Exo KM, Oosterbeek K, Lok T, Eising CM & Koffijberg K (2010) Do changes in the frequency, magnitude and timing of extreme climatic events threaten the population viability of coastal birds? Journal of Applied Ecology 47: 720-730. PDF
 
Vahl WK, Lok T, van der Meer J, Piersma T & Weissing FJ (2005) Spatial clumping of food and social dominance affect interference competition among ruddy turnstones. Behavioral Ecology 16: 834-844. PDF
 
Other publications
 
Foekema E, van der Molen J, Asjes A, Bijleveld AI, Brasseur SMJM, Camphuysen CJ, van Franeker JA, Holthuijsen S, Kentie R, Kuhn S, Leopold M, Kleine Schaars L, Lok T, Niemann H, Schop J. 2021 Ecologische effecten van het indicent met de MSC Zoe op het Nederlandse Waddengebied, met focus op microplastics. NIOZ-rapport, 2021-03. NIOZ Royal Netherlands Institute for Sea Research: Texel. 98 pp.  Link

de Goeij P, Overdijk O, Lok T & Piersma T (2015). The changing fortunes of the spoonbills of the East-Atlantic Flyway. Chapter in La Migracion Intercontinental De La Espatula (Platalea leucorodia). 

Lok T (2013) Spoonbills as a model system: a demographic cost-benefit analysis of differential migration. PhD thesis. University of Groningen. PDF
 
Lok T, Overdijk O, Horn H & Piersma T (2009) The Spoonbill Platalea leucorodia population of the Wadden Sea islands: does population growth level off? Limosa 82: 149-157. PDF